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e-mail: inmarbio@mail.primorye.ru
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GZ hohlhfhjkdhf r_evn_ Dmjbevkdbo hkljh\h\ \i_j\u_ h[gZjm`_gu q_luj_ \b^Z kde_jZdlbgbc h^bghqguo dZjbhnbeebb^ Caryophyllia ambrosia, C. scobinosa, C. alaskensis b Polymyces montereyensis GZoh^db wlbo
dhjZeeh\iha\heyxlmlhqgblvZj_ZejZg__h]jZgbq_gguc^eylj_obagboljhibq_kdbfbbl_ieufb\h^ZfbBg^hiZpbnbdb Ihemq_ggu_ k\_^_gby ihihegyl j_]bhgZevgu_ k\h^db ih nZmg_ ^hgguo khh[s_kl\ kZfhc ijh^mdlb\ghcbbgl_gkb\ghhk\Zb\Z_fhcahgur_evnZmf_j_gguo\h^K_\_jghcIZpbnbdb
Dexq_\u_keh\ZZ]_jfZlbigu_kde_jZdlbgbbDmjbevkdb_hkljh\ZjZkijhkljZg_gb_
New records of ahermatypic scleractinians from the Kuril Islands. Yu. Ya. Latypov (Institute of Marine Biology, Far East Branch, Russian Academy of Sciences, Vladivostok 690041)
Four species of solitary caryophylliid scleractinia (Caryophyllia ambrosia, C. scobinosa C. alaskensis, and Polymyces montereyensis) are described, for the first time, from the Okhotsk Sea shelf of the Kuril Islands. The findings of
these corals extend their geographic ranges, which for three of them were previously limited to tropical and warm waters of
the Indo-Pacific, and are an addition to regional check-lists of the benthic communities fauna of the most productive and intensively exploited temperate northern Pacific shelf. (Biologiya Morya, Vladivostok, 2004, vol. 30, no. 6, pp. 473–476).
Key words: Scleractinia, ahermatypic, Kuril Islands, distribution.
<i_j\u_ h^bghqgu_ Z]_jfZlbigu_1 kde_jZdlbgbb \
jhkkbckdbo \h^Zo [ueb h[gZjm`_gu gZ Z[bkkZevguo ]em[bgZo DmjbehDZfqZlkdh]h `_eh[Z D_ee_j ;ueb
gZc^_gu b hibkZgu q_luj_ \b^Z >\Z ba gbo ^hihegbeb jh^
Fungiacyathus – F. palifera b F. symmetricus Z gh\uc hibkZgguc ih^\b^ F. symmetricus symmetricus D_ee_j [ue
hlg_k_g Dwcjgkhf &DLUQV \ kbghgbfbx \b^Z F.
marenzellery>\Z^jm]bo\b^Zhlg_k_gudjh^mCaryophyllia –
C. ambrosiabC. alaskensis;he__ihegu_^Zggu_hjZaghh[jZabbZ]_jfZlbiguodhjZeeh\K_\_jghcIZpbnbdbbbojZkijhkljZg_gbb ijb\_^_gu \ ebl_jZlmj_ 9DXJKDQ 'XUKDP
1947; Durham, Barnard, 1952; Cairns, 1991).
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^h\hevgh h[uqgh_ y\e_gb_ Hgb ihklhyggh \klj_qZebkv ijb
]em[hdh\h^ghf ^jZ]bjh\Zgbb m ih[_j_`vy :eykdb Dmjbevkdbo DhfZg^hjkdbo b :e_mlkdbo hkljh\h\ 9DXJKDQ D_ee_j &DLUQV Ij_^klZ\bl_eb dZjbhnbeebb^
[he__\b^h\h[gZjm`_gugZ]em[bgZohl^hfih
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\dhlhjhcijh\_^_gZj_\babyfgh]bo\b^h\wlbodhjZeeh\
<b^C. ambrosia±i_j\ucbahibkZgguogb`_±h[blZ_l
\l_ieuo\h^ZoYihgbbZlZd`_\:leZglbd_gZ]em[bg_±
f :j_Ze \lhjh]h \b^Z C. scobinosa h]jZgbq_g ljhibq_kdbfb b l_ieufb \h^Zfb Bg^hiZpbnbdb Ij_^klZ\bl_eb
lj_lv_]h \b^Z C. alaskensis \klj_qZxlky \h fgh]bo jZchgZo
Lboh]h hd_ZgZ ± hl Dhj_ckdh]h ijheb\Z ^h :e_mlkdbo b DhfZg^hjkdbohkljh\h\Q_l\_jluc\b^Polymyces montereyensis
jZkijhkljZg_gmaZiZ^gh]hih[_j_`vyK_\_jghc:f_jbdb±hl
aZe Fhgl_j_c DZebnhjgby ^h f ;eZgdh Hj_]hg <k_ q_luj_\b^Z\i_j\u_gZc^_gugZr_evn_Dmjbevkdbohkljh\h\
khohlhfhjkdhc klhjhgu Gh\u_ gZoh^db kde_jZdlbgbc iha\heyxljZkrbjblv^Zggu_h[boZj_Ze_b]em[bg_jZkijhkljZg_gby Ihemq_ggu_ k\_^_gby iha\heyl ihihegblv j_]bhgZevgu_k\h^dbihnZmg_^hgguokhh[s_kl\kZfhcijh^mdlb\ghcb
bgl_gkb\gh hk\Zb\Z_fhc ahgu r_evnZ mf_j_gguo lbohhd_Zgkdbo\h^bamq_gghc^Ze_dhg_iheghDhjZeeugZc^_gugZ\ZemgZo ijb ^jZ]bjh\Zgbb \h \j_fy wdki_^bpbhgguo jZ[hl
\ 29fj_ck_GBK><HJ:G:dZ^_fbdHiZjbg\]
Hljy^ SCLERACTINIA
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J h ^ Caryophyllia Lamarck, 1816
Caryophyllia ambrosia Alcock, 1898
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Caryophyllia ambrosia: Alcock (1898); Zibrowius (1980);
D_ee_j
Caryophyllia cf. alcocki: Yabe, Eguchi (1942).
Caryophyllia ambrosia ambrosia: Cairns (1994).
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[hjha^dZfbihebgbyfaZdeZ^dbk_il<\_jog_cqZklbdhjZeeblZdhklZevgu_j_[jZ\ujZ`_gu[he__j_ev_ngh
Nhjfbjmxlky k_ilu iylb pbdeh\ Hgb jZg`bjh\Zgu \
ljb jZaf_jguo ihjy^dZ K_ilu i_j\h]h ihjy^dZ i_j\h]h
\lhjh]h b g_dhlhju_ lj_lv_]h pbdeh\ keb\Zxlky k dhemf_eehc Hk_\u_ djZy ba gbo hdZgqb\Zxlky ba\bebklufb \u-
:]_jfZlbigu_kde_jZdlbgbb±wlhdhjZeeug_bf_xsb_kbf[bhlbq_kdbofbdjh\h^hjhke_cahhdkZgl_ee]em[bgZbojZkijhkljZg_gbyg_
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1
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Jbk. 1. <g_rgbc \b^ qZrdb b nhjfZ dhjZeeblh\. Z, [ – Caryophyllia ambrosia (wda. 9321); \, ] – C. scobinosa (wda. 9705); ^, _ – C. alaskensis (wda. 9320). M\.: ×2.
klmiZxsbfb iZeyfb K_ilu \lhjh]h ihjy^dZ ^hoh^yl ^h dhemf_eeug_keb\Zykvkg_cK_ilulj_lv_]hihjy^dZq_l\_jlh]h b iylh]h pbdeh\ \iheh\bgm beb gZ lj_lv dhjhq_ k_il
\lhjh]h ihjy^dZ k hq_gv lhgdbfb k\h[h^gufb hk_\ufb
djZyfb;hdh\u_ih\_joghklbk_ili_j\uo^\moihjy^dh\ihdjulufgh]hqbke_ggufbrbibdZfbH[s__qbkehk_il
Dhemf_eeZ djm]eZy iehlgZy khklhbl ba keb\Zxsboky
jZaghh[jZaghbah]gmluoieZklbghd
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]em[bgZ±f
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Yihgbyie_cklhp_gHdbgZ\uYihgby±hkljh\ZOhgkxJxdx]em[bgZ±f:leZglbdZ]em[bgZ±f
Caryophyllia scobinosa Alcock, 1902
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Caryophyllia scobinosa: Alcock (1902); Yabe, Eguchi
(1942); Cairns, Keller (1993); Cairns (1994).
Caryophyllia cf. scobinosa: Utinomi (1956); Cairns (1994).
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]em[bgZf
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Yihgby ± hkljh\Z Ohgkx Jxdx Dxkx ]em[bgZ ± f
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Caryophyllia alaskensis Vaughan, 1941
Jbk^_
Caryophyllia alaskensis: Vaughan (1941); Durham (1947);
D_ee_j&DLUQVEZluih\
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i_j\h]hb\lhjh]hpbdeh\bf_xljZ\gmx^ebgmbh^bgZdh\mx
nhjfm Hgb keb\Zxlky kdhemf_eehc b bf_xl ohjhrh \ujZ`_ggu_ iZeb dhlhju_ \uklmiZxl \\_jo gZ^ ]eZ\gufb i_j_]hjh^dZfb K_ilu lj_lv_]h pbdeZ gZ lj_lv dhjhq_ ]eZ\guo
k_il Bo hk_\u_ dhgpu ijyfu_ k\h[h^gu_ hl\_kgh \_jlbdZevgu_K_iluq_l\_jlh]hpbdeZdhjhldb_g_[he__±ff
Bolhgdb_keZ[hba\bebklu_hk_\u_djZyk\h[h^gu<k_k_ilu
ihdjuluf_edbfbrbibdZfbhjb_glbjh\Zggufb\km[iZjZee_evgu_ \__jhh[jZagu_ jy^u g_fgh]h gZdehg_ggu_ d hkb
H[s__dhebq_kl\hk_il
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ba]b[Zxsbfbky mlhes_ggufb ieZklbgZfb bgh]^Z keb\Zxsbfbky^jm]k^jm]hf
F_klhgZoh`^_gb_ Dmjbevkdb_ hkljh\Z h\ Hg_dhlZg
]em[bgZ±f
JZkijhkljZg_gb_ :e_mlkdb_ DhfZg^hjkdb_ b Dmjbevkdb_ hkljh\Z Zjobi_eZ] :e_dkZg^jZ aZeb\u :eykdZ b I_ljZ
<_ebdh]hDhj_ckdbcijheb\]em[bgZ±f
AZf_qZgb_ Dmjbevkdb_ dhjZeeu ih [hevrbgkl\m ijbagZdh\koh^gukjZg__hibkZggufbwda_fieyjZfb C. alaskensis D_ee_j &DLUQV EZluih\ h^gZdh hlebqZxlky hl gbo km[pbebg^jbq_kdbfb \ukhdbfb dhjZeeblZfb
hq_gv fZe_gvdhc weebikh\b^ghc dhemf_eehc b h^bgZdh\uf
dhebq_kl\hfk_ilijbjZaguoihi_j_qguojZaf_jZoqZrdb
J h ^ Polymyces Cairns, 1979
Polymyces montereyensis (Durham, 1947)
Jbk
Flabellum montereyense: Durham (1947); Durham, Barnard
(1952).
Flabellum tannerense: Durham, Barnard (1952).
Polymyces montereyensis: Cairns (1979, 1991, 1994).
Polymyces montereyense: Bythell (1986).
Polymyces tannerensis: Cairns (1991).
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iehkdhcih^hr\hcbf_xs_ckf\^bZf_lj_QZrdZ]em[hdZy weebilbq_kdZy jZaf_jhf ± ff \ ihi_j_qgbd_ ijb
\ukhl_ dhjZeeblZ ± ff Kl_gdZ lhgdZy nZjnhjh\b^gZy
`_fqm`gh[_eh]h p\_lZ k lhgdbfb _ehqdh\b^gufb mahjZfb
DhklZevgu_j_[jZlheklu_gbadb_imgdlbjgu_
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ih ^ebg_ Hgb ihqlb ^hklb]Zxl hkb bo jh\gu_ hk_\u_ djZy
ke_]dZijbiexkgmlu_K_ilulj_lv_]hpbdeZgZ±ffdhjhq_ K_ilu q_l\_jlh]h pbdeZ \iheh\bgm beb gZ lj_lv dhjhq_
k_il lj_lv_]h pbdeZ K_ilu iylh]h pbdeZ dhjhldb_ ± ff
<k_ k_ilu bf_xl hl\_kgu_ hk_\u_ djZy bo [hdh\u_ ih\_joghklb ]jZgmebjh\Zgu hq_gv f_edbfb fgh]hqbke_ggufb rbibdZfbH[s__dhebq_kl\hk_il
Jm^bf_glZjgZy dhemf_eeZ fh`_l [ulv \ hkgh\Zgbb
qZrdbQZr_qgZyyfdZ]em[hdZyweebikh\b^gZyh]jZgbqb\Z_lky\gmlj_ggbfbdjZyfbk_ili_j\h]hb\lhjh]hpbdeh\
F_klhgZoh`^_gb_ Dmjbevkdb_ hkljh\Z h\ Hg_dhlZg
]em[bgZ±f
JZkijhkljZg_gb_ Dmjbevkdb_ hkljh\Z aZiZ^gh_ ih[_j_`v_ K_\_jghc :f_jbdb hl aZe Fhgl_j_c DZebnhjgby ^h
f;eZgdhHj_]hg]em[bgZ±f
HibkZggu_dhjZeeuojZgylky\fma__BgklblmlZ[bheh]bbfhjy><HJ:G‹
;BHEH=BYFHJYlhf‹
Jbk <g_rgbc \b^ qZrdb Z b nhjfZ dhjZeeblZ [ m
Polymyces montereyensiswdaM\×1.5.
:\lhj ijbagZl_e_g =F DZf_g_\m aZ ij_^hklZ\e_gguc
fZl_jbZebK>Kl_iZgvygpaZp_ggu_aZf_qZgbydjmdhibkb
klZlvb
KIBKHDEBL?J:LMJU
D_ee_j G; =em[hdh\h^gu_ fZ^j_ihjh\u_ dhjZeeu jh^Z
FungiaciathusbaDmjbehDZfqZlkdh]hb:e_mlkdh]h`_eh[h\bg_dhlhjuo^jm]boj_]bhgh\Fbjh\h]hhd_ZgZ
LjBH:GKKKJLK±
D_ee_j G; F_`\b^h\Zy baf_gqb\hklv Caryophyllia
0DGUHSRUDULD \ k\yab kbo mkeh\byfb h[blZgby Lj
BH:GKKKJLK±
EZluih\ XY Gh\u_ k\_^_gby h kde_jZdlbgbyo ba k_\_jh
aZiZ^ghc qZklb Yihgkdh]h fhjy ;bhe fhjy L‹K±
Alcock A. An account of the deep-sea Madreporaria collected by
the royal Indian marine survey ship "Investigator". Calcutta:
Trustees of the Indian Museum. 1898. 29 p.
Alcock A. Diagnoses and description of new species of corals //
Tijdschr. Neder. Dierkund. Vereenin. 1902. Vol. 2, no. 7.
P. 89–115.
Bythell J.C. A guide to the identification of the living corals
(Scleractinia) of Southern California // Occas. Pap. San Diego Soc. Nat. Hist. 1986. Vol. 16. 40 p.
Cairns S.D. The deep-water Scleractinia of the Caribbean Sea and
adjacent waters // Studies on the fauna of Curaçao and other
Caribbean Islands. 1979. Vol. 180. 341 p.
K?EBG<?OH<:
476
Cairns S.D. A revision of the ahermatypic Scleractinia of the Galapagos and Cocos Islands // Smithsonian Contrib. Zool.
1991. No. 504. 44 p.
Cairns S.D. Scleractinia of the temperate North Pacific // Smithsonian Contrib. Zool. 1994. No. 557. 150 p.
Cairns S.D., Keller N.B. New taxa and distributional records of
azooxanthellate Scleractinia (Cnidaria: Anthozoa) from the
tropical southwest Indian Ocean, with comments on their
biogeography and biology // Ann. S. Afr. Mus. 1993.
Vol. 103, no. 5. P. 213–292.
Durham J.W. Corals from the Gulf of California and the North
Pacific coast of America // Mem. Geol. Soc. Amer. 1947.
Vol. 20. 68 p.
Durham J.W., Barnard J.L. Stony corals of the Eastern Pacific
collected by the Velero III and Velero IV // Allan Hancock
Pac. Exp. 1952. Vol. 16. 110 p.
Utinomi H. Invertebrate fauna of the intertidal zone of the Tokara
Islands, XIV: stony corals and hydrocorals // Bull. Mun.
Mus. Nat. Hist. 1956. Vol. 10. P. 37–44.
Vaughan T.W. New corals: one recent., Alaska; three Eocene,
Alabama and Louisiana // J. Paleont. 1941. Vol. 15, no. 3.
P. 280–284.
Yabe H., Eguchi M. Fossil and recent simple corals from Japan //
Sci. Rept. Tohoku Imper. Univ. Ser. 2 (Geology). 1942.
Vol. 22, no. 2. P. 105–178.
Zibrowius H. Les Scléractiniares de la Méditeranée et de Atlantique Nord-Oriental // Mem. Inst. Océanogr. Monaco. 1980.
Vol. 11. 284 p.
;BHEH=BYFHJYlhf‹k±478
M>D
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>BG:FBD:H;J:AH<:GBY;BKKMKGUOGBL?C
M CRENOMYTILUS GRAYANUSBMODIOLUS MODIOLUS (BIVALVIA)
IJBIH<LHJGHFIJBDJ?IE?GBBDKM;KLJ:LM
‹]GBK_ebg??<_oh\Z
Bgklblml[bheh]bbfhjy><HJ:G<eZ^b\hklhd
e-mail: inmarbio@mail.primorye.ru
KlZlvyijbgylZdi_qZlb]
MklZgh\e_ghqlhijhp_kkih\lhjgh]hijbdj_ie_gbydkm[kljZlmm^\mkl\hjqZluofheexkdh\Crenomytilus
grayanusbModiolus modioluskeZ]Z_lkybag_kdhevdboihke_^h\Zl_evguowlZih\aZgbfZxsbo\wdki_jbf_glZevguomkeh\byoijbl_fi_jZlmj_\h^u± 2ºKhdhehh^gh]hf_kypZIhdZaZghqlhfh^bhemkbf__l[he__\ukhdb_
l_fiuh[jZah\Zgby[bkkmkguogbl_cq_ffb^by=j_yZlZd`_[hevr__bodhebq_kl\hddhgpmijhp_kkZnhjfbjh\Zgby iheghp_ggh]h [bkkmkgh]h dhfie_dkZ GZ[ex^Z_fu_ jZaebqby h[tykgyxlky Z^ZilZpb_c fheexkdh\ d
h[blZgbx\jZaguo[bhlhiZo
Dexq_\u_keh\Z[bkkmkgu_gblbih\lhjgh_ijbdj_ie_gb_Crenomytilus grayanus, Modiolus modiolus.
Dynamics of byssus thread formation in Crenomytilus grayanus and Modiolus modiolus (Bivalvia) at reattachment to substrate. N. I. Selin, E. E. Vekhova (Institute of Marine Biology, Far East Branch, Russian Academy of
Sciences, Vladivostok 690041)
Reattachment process in the bivalves Crenomytilus grayanus and Modiolus modiolus involves several successive
stages, which take in vitro about one month at 19 ± 2ºC. In comparison with C. grayanus, M. modiolus had higher rates
of byssus thread production and a larger number of byssal threads by the end of the byssus complex formation. The differences can be explained by the adaptation of mussels for life in different biotopes. (Biologiya Morya, Vladivostok,
2004, vol. 30, no. 6, pp. 476–478).
Key words: byssus threads, reattachment, Crenomytilus grayanus, Modiolus modiolus.
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