НОВЫЕ ВИДЫ МОРСКИХ СТРЕЛОК (CHAETOGNATHA) ИЗ
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;BHEH=BYFHJYlhfk440–446 M>D AHHEH=BY;?KIHA<HGHQGUO GH<U?<B>UFHJKDBOKLJ?EHDCHAETOGNATHA) BA;MOLUQ:@F:A:EB<:I?LJ:<?EBDH=H YIHGKDH?FHJ?1 ]:IDZkZldbgZ:NK_j]__\ Lbohhd_Zgkdbchd_Zgheh]bq_kdbcbgklblml><HJ:G<eZ^b\hklhd _-mail: pacific@online.marine.su KlZlvyijbgylZdi_qZlb] <[QZ`fZaZeI_ljZ<_ebdh]hYihgkdh]hfhjyh[gZjm`_gh\b^h\fhjkdboklj_ehdbadhlhjuo\b^Z (Leptosagitta ocis sp. n., L. exigua sp. n., L. icis sp. nbOligoradiata entis sp. nhibkZgudZdgh\u_^eygZmdbG_dhlhju_wda_fieyjubf_ebg_h[uqgu_fhjnheh]bq_kdb_ZghfZebbdhlhju_g_[uebhlf_q_gumfhjkdboklj_ehd ba\medZgbq_kdbo[molb]_hnbabq_kdbZdlb\guoj_]bhgh\<klZlv_ZgZebabjm_lkywdheh]bq_kdZyh[klZgh\dZ\ [QZ`fZ ijb\_^_gu kibkhd h[gZjm`_gguo \b^h\ fhjkdbo klj_ehd k hibkZgb_f b jbkmgdZfb gh\uo \b^h\ Z lZd`_i_j_q_gvfhjnheh]bq_kdbohldehg_gbchlf_q_gguom\k_o\b^h\klj_ehd Dexq_\u_keh\Zfhjkdb_klj_edbChaetognathaieZgdlhgfhjnheh]bq_kdb_hldehg_gbyjZ^bhZdlb\gh_ aZ]jyag_gb_ New species of arrowworms (Chaetognatha) from Chazhma Bay (Peter the Great Bay, Sea of Japan). A. P. Kasatkina, A. F. Sergeev (Pacific Oceanological Institute, Far East Branch, Russian Academy of Sciences, Vladivostok 690041) Eighteen species of Chaetognatha were discovered in Chazhma Bay (Peter the Great Bay, Sea of Japan). Of these, 4 new species (Leptosagitta ocis sp. n., L. exigua sp. n., L. icis sp. n., and Oligoradiata entis sp. n.) are described as new for science. Some of specimens had morphological abnormalities, which were not observed in chaetognaths from volcanic bays and active geophysical regions. The ecological situation in Chazhma Bay is analyzed. A list of chaetognaths along with descriptions and figures of new species are provided. Morphological abnormalities found in all species of chaetognaths are outlined. (Biologiya Morya, Vladivostok, 2004, vol. 30, no. 6, pp. 440–446). Key words: arrowworms, Chaetognatha, plankton, morphological abnormalities, radioactive contamination. hd_Zgheh]bq_kdhf bgklblml_ ><H J:G ] <eZ^b\hklhd \ lbih\hcfZl_jbZeg_\dexq_guhkh[bkZghfZebyfb < i_jbh^ hl[hjZ ieZgdlhgguo ijh[ \ [ QZ`fZ gZfb ijh\h^bebkv jZ^bhwdheh]bq_kdb_ bkke_^h\Zgby g_dhlhju_ j_amevlZlu dhlhjuo gZjy^m kebl_jZlmjgufb ^Zggufb bkihevah\Zgu^eyijb\h^bfh]hgb`_djZldh]hhibkZgbyjZchgZ jZ[hl ; QZ`fZ jZkiheh`_gZ \ k_\_jh\hklhqghc qZklb aZe I_ljZ<_ebdh]h>gh[molukeh`_ghh^ghjh^gufbih]jZgmehf_ljbq_kdhfmkhklZ\mhkZ^dZfbkij_h[eZ^Zgb_ffbdlblh\ Ze_\jblh\uo b i_kqZguo :klZoh\ bgh]^Z k aZiZohf k_jh\h^hjh^Z Abfhc [molZ dZd ijZ\beh aZf_jaZ_l E_lhf \h^Z\ih\_joghklghfkeh_ijh]j_\Z_lky^hºKb\ur_OZjZdl_jgu_^ey\h^[moluagZq_gbykhe_ghklbe_`Zl\^bZiZahg_ ±Å < dmlh\hc qZklb [molu khe_ghklv \h^u gb`_ \ke_^kl\b_ jZkij_kgyxs_]h \ebygby \iZ^Zxs_c \ g__ jX`ZgdZ < p_ehf [ QZ`fZ oZjZdl_jbam_lky keZ[hc ^bgZfbdhc\h^bbf__l^\mokehcgmxdhfi_gkZpbhggmxkljmdlmjm pbjdmeypbb Kdhjhklv \ih\_joghklghf keh_ h[uqgh g_ ij_\urZ_l±kfkm^gZ±±kfk6HUJHHYHWDO :d\Zlhjby [ QZ`fZ y\ey_lky qZklvx km^hj_fhglgh]h aZ\h^ZkieZ\mqbfbibjkZfb^hdZfbb[Zabjmxsbfbkykm^ZfbHgZoZjZdl_jbam_lky\ukhdbfmjh\g_faZ]jyag_gbyk\hckl\_gguf ih^h[ghfm lbim [mol Hij_^_e_gb_ kh^_j`Zgby ly`_euof_lZeeh\\\h^_^hgguohkZ^dZob\h\a\_kbk\b^_l_evkl\m_lhlhfqlhbomjh\_gv\[QZ`fZebrvg_kdhevdh gb`_q_f\[AhehlhcJh]baZeGZoh^dZdhlhju_y\eyxlky Ijb[j_`gu_\h^ux`gh]hIjbfhjvyih^\_j`_gu bgl_gkb\ghfmZgljhih]_gghfmbl_ogh]_gghfm\ha^_ckl\bx qlh ijb\_eh d baf_g_gbyf \wdhkbkl_fZo b d kgb`_gbx bo [bhijh^mdlb\ghklb < k\yab k wlbf bamq_gb_ khklhygby ieZgdlhgguo wdhkbkl_f b \uy\e_gb_ wdheh]bq_kdboihke_^kl\bcaZ]jyag_gby±h^gZba\Z`guo aZ^Zq ijb bkke_^h\Zgbb [bhjZaghh[jZaby b ijh^mdlb\ghklbfhjkdbo[bhj_kmjkh\j_]bhgZ < ^Zgghc jZ[hl_ ijb\_^_gu j_amevlZlu bamq_gby \b^h\h]h khklZ\Z o_lh]gZl &KDHWRJQDWKD b bo fhjnheh]bq_kdbo hldehg_gbc \ ieZgdlhg_ [QZ`fZ ih^\_j`_gghc dZd h[uqghfm aZ]jyag_gbx k\hckl\_gghfm ^Zgghfmlbim[mollZdbZ\ZjbcghfmjZ^bhZdlb\ghfm ; QZ`fZ y\ey_lky lZd`_ bklhqgbdhf aZ]jyag_gby ijbe_]Zxs_]hdg_caZe Klj_ehd [ JZa[hcgbd qlh ijb\_eh d kgb`_gbx [bhfZkku b \b^h\h]h jZaghh[jZaby [_glhkZ\wlhfaZeb\_;_eZg F:L?JB:EBF?LH>BD: Ijh[uieZgdlhgZhlh[jZgu:NK_j]__\uf\[QZ`fZ \ ± ]] k_lvx >`_^b ^bZf_lj \oh^gh]h hl\_jklby 37ff hl ^gZ ± f ^h ih\_joghklb Ijh[u nbdkbjh\Zeb nhjfZev^_]b^hfaZl_fZgZebabjh\Zebih^[bghdmeyjhf Lbiu b iZjZlbiu gh\uo \b^h\ ojZgylky \Lbohhd_Zgkdhf 1 JZ[hlZ \uiheg_gZ ijb nbgZgkh\hc ih^^_j`d_ ]jZglh\ Dhgdmjk ><H J:G ] : ? NPIBgl_]jZpbyW 440 GH<U?<B>U 441 gZb[he__aZ]jyag_ggufbZd\ZlhjbyfbaZeI_ljZ<_ebdh]h< [ QZ`fZ b gZ ijbe_]Zxs_c Zd\Zlhjbb lZd`_ \ukhd mjh\_gv aZ]jyag_gby g_nl_ijh^mdlZfb K_j]__\ b ^j Hlebqbl_evghc wdheh]bq_kdhc hkh[_gghklvx wlhc [molu y\ey_lky jZ^bhZdlb\gh_ aZ]jyag_gb_ __ ^hgguo hkZ^dh\ dhlhjh_ ijhbahreh\ke_^kl\b_Z\ZjbbZlhfghcwg_j]_lbq_kdhcmklZgh\db ih^\h^ghc eh^db \ ] GZb[he__ aZ]jyag_gghc hdZaZeZkv x]haZiZ^gZy qZklv [molu ]^_ hl ahgu wibp_gljZ Z\Zjbb \ ^hgguo hkZ^dZo \^hev ih[_j_`vy knhjfbjh\Zeky Zkbff_ljbqgucjZ^bhZdlb\gucke_^Hkgh\gufdhfihg_glhfjZ^bhZdlb\gh]haZ]jyag_gbyy\ey_lkydhjjhabhggucKh> 99%) b qZklbqgh ± ijh^mdl y^_jgh]h ^_e_gby &V KhklZ\ hkgh\guoaZ]jyagbl_e_ch[mkeh\e_gl_fqlhZ\ZjbyijhbahreZ k j_Zdlhjhf aZiheg_gguf k\_`bf y^_jguf lhieb\hf Kb\bgp_\b^j<gZklhys__\j_fyfZdkbfZevgZym^_evgZyZdlb\ghklvKhb&V\^hgguohkZ^dZo\ahg_Zd\Zlhjbb\k__s_\_ebdZbij_\urZ_l7b3;dd]khhl\_lkl\_gghHq_\b^ghqlhijb^hgguckehc\h^uih^\_j]Z_lkyh[emq_gbx DZd ihdZaZeb jZ^bhf_ljbq_kdb_ baf_j_gby bgl_gkb\ghklv]ZffZbaemq_gby\g_dhlhjuof_klZoahguZ\Zjbb\ \h^ghclhes_kgb`Z_lky^hnhgh\h]hagZq_gbylhevdhgZjZkklhygbb f hl ^gZ LZdbf h[jZahf \ ahg_ Z\Zjbb fhjkdb_ hj]Zgbafu gZoh^ysb_ky \ ijb^hgghf keh_ \h^ghc lhesb b gZ^g_ZlZd`_\lhes_hkZ^dh\bkiulu\Zxl^hihegbl_evgmxdijbjh^ghcjZ^bZpbhggmxgZ]jmadm J?AMEVL:LUBH;KM@>?GB? BaieZgdlhgguoijh[[ueh\u[jZghbh[ke_^h\Zgh k\ur_ wda fhjkdbo klj_ehd Ba h[gZjm`_gguo\b^h\fhjkdboklj_ehdhdZaZebkvba\_klgufb \b^Zfb DZkZldbgZ DZkZldbgZ IZgl_e__\Z DZkZldbgZK_eb\Zgh\ZZ[Bajh^ZLeptosagitta h[gZjm`_gh iylv ba\_klguo (L. nudata Kassatkina, 1973; L. collariata Kassatkina, 1973; L. schakovi Kassatkina, 1973; L. eris Kassatkina et Selivanova, 2003; L. alba .DVVDWNLQD HW6HOLYDQRYD b ljb gh\uo \b^Z (L. icis sp. n., L. exigua VSQbL. ocis sp. n.)bajh^Z Aidanosagitta ± k_fv ba\_klguo \b^h\ A. scarlatoi Kassatkina, 1971; A. macilenta Kassatkina, 1971; A. acus Kassatkina et Selivanova, 2003; A. pilum Kassatkina et Selivanova, 2003; A. murex Kassatkina et Selivanova, 2003; A. bella Kassatkina et Selivanova, 2003; A. venusta Kassatkina et SelivanoYD ba jh^Z Parasagitta ± h^bg ba\_klguc \b^ (P. septicoela.DVVDWNLQDZlZd`_h[gZjm`_guh^bg gh\uc\b^bajh^ZOligoradiata (O. entis VSQbh^bgwda_fieyjbagh\h]hjh^ZhibkZgb_dhlhjh]hg_ijb\h^blkylZddZdwda_fieyjgZoh^bekygZcklZ^bbaj_ehklb G_h[oh^bfh hlf_lblv qlh ^ey hibkZgby gh\uo \b^h\ bkihevah\Zeb wda_fieyju g_ bf_xsb_ fhjnheh]bq_kdbohldehg_gbc Leptosagitta icis sp. n. Jbk:= >bZ]ghaI_j_^gbcdhg_pieZ\gbdh\ciZjugZqbgZ_lky \i_j_^b mjh\gy aZ^g_]h dhgpZ [jxrgh]h ]Zg]eby wlZ qZklv ieZ\gbdZ e_`Zs_]h gZ mjh\g_ ]Zg]eby dhjhq_ ]Zg]eby \ jZa Emqb g_ihegu_ lhevdh \ [hdh\uo ieZ\gbdZo ih bo djZyf bf_xlky [_aemq_\u_ ahgu K_f_ggu_ f_rhqdb khijbdZkZxlky dZd k [hdh\ufblZdbko\hklh\ufieZ\gbdZfb ;BHEH=BY FHJY lhf 30 6 2004 Jbk. 1. =hehlbiu. :, = – Leptosagitta icis sp. n. (: – h[sbc \b^, = – ]eZa); ;, < – L. exigua sp. n. (; – h[sbc \b^, < ±]eZa FZl_jbZe =hehlbi Bg\_glZjguc ghf_j /3 H[gZjm`_g\[QZ`fZaZeI_ljZ<_ebdh]hYihgkdh]h fhjyfZjlZ]ijbhljbpZl_evghcl_fi_jZlmj_ IZjZlbiu Q_luj_ wda_fieyjZ gZc^_gu \ lhc `_ ijh[_ qlh b ]hehlbi Bg\_glZjgu_ ghf_jZ /3 LP/3b/3 D:K:LDBG:K?J=??< 442 HibkZgb_ ]hehlbiZ L_eh jb]b^gh_ fmkdmebklh_ =heh\Zke_]dZrbj_lmeh\bsZgZlmeh\bsgho\hklh\hc i_j_]hjh^d_km`_gbyg_lWda_fieyjgZcklZ^bbaj_ehklb>ebgZl_eZff>ebgZo\hklh\h]hhl^_eZkhklZ\ey_lZ[jxrgh]h]Zg]ebyhl^ebgul_eZ <ujhklulmeh\bsghcdbrdbhlkmlkl\mxlLmeh\bsgZy dbrdZg_\Zdmhebabjh\ZgZghkiexs_gZ\^hjkh\_gl jZevghf gZijZ\e_gbb F_jpZl_evgZy i_ley dhjhldZy gZqbgZ_lkyihaZ^bmjh\gy]eZaqmlv\ur_r_bg_bf__l gb h^ghc iZju f_^bZevguo eZdmg I_j_^gbc dhg_p ieZ\gbdh\ciZjugZqbgZ_lky\i_j_^bmjh\gyaZ^g_]h dhgpZ[jxrgh]h]Zg]ebywlZqZklvieZ\gbdZe_`Zs_]h gZ mjh\g_ ]Zg]eby dhjhq_ ]Zg]eby \ jZa IeZ\gbdb 1ciZjudhjhq_[hdh\uoieZ\gbdh\ciZjubo^ebgZ khklZ\ey_lhl^ebgul_eZI_j_^gbcdhg_pieZ\gbdh\ciZjuhlklhblhlaZ^g_]hdhgpZieZ\gbdh\c iZju gZ jZkklhygb_ khklZ\eyxs__ ijbf_jgh hl ^ebgul_eZLmeh\bsgZyqZklvieZ\gbdh\ciZjudhjhq_boo\hklh\hcqZklb\jZaZO\hklh\hcieZ\gbd aZdjm]e_gguc_]h^ebgZgZl_e_^hklb]Z_lhlh[s_c ^ebgubhl^ebguo\hklh\h]hhl^_eZEmqbg_ihegu_lhevdh\[hdh\uoieZ\gbdZoihbodjZyfbf_xlky [_aemq_\u_ ahgu :ev\_heyjgZy ldZgv lhgdbf keh_f ihdju\Z_l \k_ l_eh ke_]dZ mlhesZykv gZ ^hjkZevghc klhjhg_gZijhlb\[jxrgh]h]Zg]ebyS_lbghdih± i_j_^gboam[qbdh\ih±aZ^gboam[qbdh\ih±\ dZ`^hfjy^m=eZaZkl_fgufib]f_glgufiylghfijyfhm]hevghc nhjfu \ p_gljZevghc qZklb Ib]f_gl aZgbfZ_lqZklvh[t_fZ]eZaZK_f_ggu_f_rhqdbm^ebg_ggu_khklZ\eyxlhdhehhl^ebgul_eZbhdheh hl ^ebgu o\hklh\h]h hl^_eZ hgb gZe_]Zxl gZ [hdh\u_ b o\hklh\hc ieZ\gbdb Ybqgbdb g_fgh]h g_ ^hoh^yl^hmjh\gyi_j_^g_]hdhgpZieZ\gbdh\ciZju Baf_gqb\hklv >ebgZ iheh\haj_euo hkh[_c dhe_[e_lkyhl^hffZhlghkbl_evgZy^ebgZo\hklh\h]hhl^_eZfh`_l^hklb]Zlv±hl^ebgul_eZ >bnn_j_gpbZevguc ^bZ]gha <b^ ohjhrh hlebqbf hl \k_o ^jm]bo \b^h\ ^Zggh]h jh^Z iheh`_gb_f ieZ\gbdh\ c iZju m \k_o hklZevguo \b^h\ hgb jZkiheh`_gu gZ mjh\g_ beb ihaZ^b mjh\gy aZ^g_]h dhgpZ [jxrgh]h]Zg]eby K\_^_gbyih[bheh]bbIhkhklhygbx]hgZ^fh`gh dhk\_ggh km^blv hjZafgh`_gbb hgh ijhbkoh^bl hq_\b^gh ijb hljbpZl_evghc l_fi_jZlmj_ \^_dZ[j_ Iheh\haj_euowda_fieyjh\\_kghcg_hlf_q_gh Wlbfheh]by. "Icis" ±b]hevqZlh_ihnhjf_h[jZah\Zgb_eZl Leptosagitta exigua sp. n. Jbk;< >bZ]ghaI_j_^gbcdhg_pieZ\gbdh\ciZjugZqbgZ_lkyihaZ^bmjh\gyaZ^g_]hdhgpZ[jxrgh]h]Zg]ebyEmqb\ieZ\gbdZoihegu_[_aemq_\ZyahgZ\h\k_o ieZ\gbdZo hlkmlkl\m_l K_f_ggu_ f_rhqdb khijbdZkZxlky dZd k [hdh\ufb lZd b k o\hklh\uf ieZ\gbdZfb Ijhf_`mlhd f_`^m ieZ\gbdZfb c b c iZju ^ebgg__ q_f f_`^m [jxrguf ]Zg]eb_f b ieZ\gbdZfb c iZju ;jxrghc ]Zg]ebc dhjhq_ lmeh\bsghc qZklb ieZ\gbdZciZju FZl_jbZe =hehlbi Bg\_glZjguc ghf_j /3 H[gZjm`_g\[QZ`fZaZeI_ljZ<_ebdh]hYihgkdh]h fhjy^_dZ[jy]ijbhljbpZl_evghcl_fi_jZlmj_ IZjZlbiu Ljb wda_fieyjZ \u^_e_gu ba lhc `_ ijh[u qlh b ]hehlbi Bg\_glZjgu_ ghf_jZ /3 LPb/P-7/4. 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"Exigua"±f_edZyeZl ;BHEH=BY FHJY lhf 30 6 2004 GH<U?<B>U 443 Leptosagitta ocis sp. n. Jbk:= >bZ]ghaI_j_^gbcdhg_pieZ\gbdh\ciZjugZqbgZ_lkyihaZ^bmjh\gyaZ^g_]hdhgpZ[jxrgh]h]Zg]eby ;hdh\u_ ieZ\gbdb \ bo i_j_^g_c qZklb ebr_gu emq_c[_aemq_\ZyahgZbf__lkylhevdh\[hdh\uoieZ\gbdZo Ijhf_`mlhd f_`^m ieZ\gbdZfb c b c iZju jZ\_g ijhf_`mldm f_`^m [jxrguf ]Zg]eb_f b ieZ\gbdZfbciZju;jxrghc]Zg]ebc^ebgg__lmeh\bsghcqZklbieZ\gbdZciZju FZl_jbZe =hehlbi Bg\_glZjguc ghf_j /3 H[gZjm`_g\[QZ`fZaZeI_ljZ<_ebdh]hYihgkdh]h fhjyn_\jZey]\ih\_joghklghfkeh_\h^uijb hljbpZl_evghcl_fi_jZlmj_ IZjZlbiu Ljb wda_fieyjZ \u^_e_gu ba lhc `_ ijh[u qlh b ]hehlbi Bg\_glZjgu_ ghf_jZ /3 LP-8/3, LP-8/4. HibkZgb_ ]hehlbiZ L_eh jb]b^gh_ fmkdmebklh_ =heh\Z rbj_ lmeh\bsZ gZ lmeh\bsgho\hklh\hc i_j_]hjh^d_ km`_gby g_l Wda_fieyj gZ c klZ^bb aj_ehklb >ebgZ l_eZ ff >ebgZ o\hklh\h]h hl^_eZ khklZ\ey_lZ[jxrgh]h]Zg]ebyhdhehhl^ebgul_eZ<ujhklulmeh\bsghcdbrdbhlkmlkl\mxlLmeh\bsgZy dbrdZ g_ \Zdmhebabjh\ZgZ g_ kiexs_gZ \^hjkh\_gljZevghf gZijZ\e_gbb gh g_ ijyfZy dZd m \k_ohklZevguo\b^h\jh^ZZba\bebklZyF_jpZl_evgZy i_ley dhjhldZy gZqbgZ_lky kjZam ihaZ^b mjh\gy ]eZa qmlv \ur_ r_b b jZkiheZ]Z_lky ihjh\gm gZ ]heh\_ b lmeh\bs_ I_ley bf__l h^gm iZjm keZ[h \ujZ`_gguo f_^bZevguo \uimdehkl_c I_j_^gbc dhg_p ieZ\gbdh\ c iZju gZqbgZ_lky ihaZ^b mjh\gy aZ^g_]h dhgpZ [jxrgh]h ]Zg]eby gZ jZkklhygbb \ jZaZ dhjhq_ [jxrgh]h]Zg]ebyIeZ\gbdbciZjudhjhq_ieZ\gbdh\ c iZju ijb[ebabl_evgh \ jZaZ b khklZ\eyxl ijbf_jghhl^ebgul_eZI_j_^gbcdhg_pieZ\gbdh\ c iZju hlklhbl hl aZ^g_]h dhgpZ ieZ\gbdh\ c iZjugZjZkklhygb_khklZ\eyxs__hdhehhl^ebgu l_eZhghjZ\ghjZkklhygbxf_`^m[jxrguf]Zg]eb_f bieZ\gbdZfbciZju>ebgZieZ\gbdh\ciZjukhklZ\ey_l [he__ hl ^ebgu l_eZ bo lmeh\bsgZy qZklv dhjhq_ o\hklh\hc \ jZaZ O\hklh\hc ieZ\gbd aZdjm]e_gguc _]h ^ebgZ gZ l_e_ khklZ\ey_l hl h[s_c^ebgubijb[ebabl_evghhl^ebguo\hklh\h]h hl^_eZ Emqb g_ihegu_ lhevdh \ [hdh\uo ieZ\gbdZo i_j_^gyyqZklvdhlhjuoiheghklvxebr_gZemq_c:ev\_heyjgZyldZgvihdju\Z_l\k_l_ehhlr_b^hdZm^Zevgh]hdhgpZmlhesZykvk^hjkZevghcklhjhgugZijhlb\ [jxrgh]h ]Zg]eby b gZ lmeh\bs_ f_`^m ieZ\gbdZfb 1cbciZjuGZ]heh\_h^gZiZjZjy^h\s_lbghd^\_ iZju jy^h\ am[qbdh\ S_lbghd ih i_j_^gbo am[qbdh\ihaZ^gboam[qbdh\ih\dZ`^hfjy^m=eZaZk l_fgufijyfhm]hevgufib]f_glhf\p_gljZevghcqZklb K_f_ggu_ f_rhqdb m^ebg_ggu_ khklZ\eyxl hdheh hl ^ebgu l_eZ b hl ^ebgu o\hklh\h]h hl^_eZ hgbkhijbdZkZxlkydZdk[hdh\ufblZdbko\hklh\uf ieZ\gbdhf Ybqgbdb ^hklb]Zxl mjh\gy i_j_^g_]h dhgpZieZ\gbdh\ciZju ;BHEH=BY FHJY lhf 30 6 2004 Jbk =hehlbiu : = ± Leptosagitta ocis sp. n : ± h[sbc \b^=±]eZa; < – Oligoradiata entis sp. n;±h[sbc\b^ <±]eZa D:K:LDBG:K?J=??< 444 Baf_gqb\hklv >ebgZ iheh\haj_euo hkh[_c dhe_[e_lky hl ^h ff Z hlghkbl_evgZy ^ebgZ o\hklh\h]h hl^_eZ fh`_l khklZ\eylv ± hl ^ebgu l_eZ[jxrgh]h]Zg]eby±±hl^ebgul_eZ >bnn_j_gpbZevguc ^bZ]gha <b^ [ebahd d L. exigua VSQghhlebqZ_lkyhlg_]hjy^hfijbagZdh\kj_^bgbogZb[he__aZf_lgu_gZebqb_[_aemq_\uoahgm L. ocis VSQjZkiheh`_gb_ieZ\gbdh\m L. ocis sp. n. ijhf_`mlhd f_`^m ieZ\gbdZfb c b c iZju jZ\_g jZkklhygbx f_`^m [jxrguf ]Zg]eb_f b ieZ\gbdZfb 1ciZjuZmL. exigua VSQhgh^ebgg__[jxrghc ]Zg]ebc m L. exigua VS Q dhjhq_ lmeh\bsghc qZklb ieZ\gbdh\ciZjumL. ocis VSQ±^ebgg__ K\_^_gbyih[bheh]bbIhkhklhygbx]hgZ^fh`gh dhk\_ggh km^blv hjZafgh`_gbb hgh ijhbkoh^bl hq_\b^gh ijb hljbpZl_evghc l_fi_jZlmj_ \^_dZ[j_ Iheh\haj_euowda_fieyjh\\_kghcg_hlf_q_gh Wlbfheh]by. "Ocis"±[ukljuceZl Oligoradiata entis sp. n. Jbk;< >bZ]gha K_f_ggu_ f_rhqdb m^Ze_gu dZd hl [hdh\uolZdbhlo\hklh\h]hieZ\gbdZgZjZ\gh_jZkklhygb_ Lmeh\bsgZy qZklv ieZ\gbdh\ c iZju agZqbl_evghdhjhq_boo\hklh\hcqZklbijbf_jgh\jZaZ FZl_jbZe=hehlbiBg\_glZjgucghf_j2/5 H[gZjm`_g \ [QZ`fZ aZe I_ljZ <_ebdh]h Yihgkdh]h fhjyn_\jZey] IZjZlbiu >\Z wda_fieyjZ \u^_e_gu ba lhc `_ ijh[u qlh b ]hehlbi Bg\_glZjgu_ ghf_jZ 2/5 OLR-1/14. 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"Entis"±ijhk\_qb\ZxsZyeZl HibkZgb_fhjnheh]bq_kdbohldehg_gbc mfhjkdboklj_ehd\[QZ`fZ G_h[oh^bfh hlf_lblv qlh i_j_qbke_ggu_ gb`_ fhjnheh]bq_kdb_ hldehg_gby g_ y\eyxlky f_oZgbq_kdbfb ih\j_`^_gbyfb Fgh]he_lgbc hiul jZ[hlu Z\lhjh\ klZlvb m[_^bl_evgh ihdZaZe qlh f_oZgbq_kdb_ ih\j_`^_gby m fhjkdbo klj_ehd fh]ml ihy\blvky \ j_amevlZl_ g_ hl[hjZ Z jZa[hjZ ijh[ g_hiulgufb ex^vfb dhlhju_ ihevamykv ibgp_lhf k ijyfufb dhgpZfb kbevghk`bfZxl`b\hlgh_ihi_j_dlmeh\bsZFujZa[bjZeb ijh[u ibgp_lhf k aZ]gmlufb dhgpZfb beb k ihfhsvxibi_ldbIjbf_qZl_evghqlhbmZghfZevguo hkh[_c hlh[jZgguo ba [ QZ`fZ b m hkh[_c ba ]_hnbabq_kdb Zdlb\guo j_]bhgh\ hklZ_lky g_ih\j_`^_gguf gZb[he__g_`gucbmya\bfuchj]Zgfhjkdboklj_ehd± f_jpZl_evgZy i_ley dhlhjZy mljZqb\Z_lky ijb g_ZddmjZlghff_oZgbq_kdhfi_j_f_s_gbbwda_fieyjh\ Gb \ h^ghf ba j_]bhgh\ h[jZ[hlZgu lukyqb ijh[±hl:leZglbdb^h<hklhqghDblZckdh]hfhjylZ;BHEH=BY FHJY lhf 30 6 2004 GH<U?<B>U dbo fhjnheh]bq_kdbo hldehg_gbc m fhjkdbo klj_ehd dZd\[QZ`fZg_\klj_qZehkvH^gZdhmfhjkdboklj_ehd ba [QZ`fZ g_h[gZjm`_gufhjnheh]bq_kdb_ ZghfZebboZjZdl_jgu_^ey`b\hlguoh[blZxsbo\\h^Zo \medZgbq_kdbo[molbk_ckfbq_kdbZdlb\guoj_]bhgh\ baf_g_gby \ ib]f_glghf gmde_Zjghf y^j_ Z bgh]^Z b ihegh_ bkq_agh\_gb_ ]eZaZ Z lZd`_ ebabk furp DZkZldbgZ.DVVDWNLQDHWDO GZb[he__ qZklh \klj_qZxsb_ky ZghfZevgu_ hldehg_gbymfhjkdboklj_ehd JZajmr_gb_ ieZ\gbdh\hc ieZklbgdb ih djZyf f_`^memq_cIjbq_femqbhklZxlkyg_ih\j_`^_ggufb bqZs_\k_]hbodjZylhjqZlaZij_^_eZfbieZ\gbdh\uo ieZklbghd jbk : Bgh]^Z \ j_amevlZl_ jZajmr_gby ieZklbgdbdjZyieZ\gbdh\g_bf_xl[ZojhfqZlhcdZ_fdbbak\h[h^guoemq_cdZdgZjbk:Z\u]ey^yldZd i_lmrbgu_]j_[gbjbk;G_h[oh^bfhhlf_lblvqlh ijbf_oZgbq_kdhfih\j_`^_gbbjZajmrZ_lkyg_lhevdh djZ_\Zyf_f[jZgZZbgh]^Zb\kyieZ\gbdh\ZyieZklbgdZghih\j_`^ZxlkybieZ\gbdh\u_emqbihwlhfmih^h[ghc dZjlbgu mf_oZgbq_kdb ih\j_`^_gguo hkh[_c g_ gZ[ex^Z_lky >ey ZghfZevguo wda_fieyjh\ ba [ QZ`fZi_lmrbguc]j_[_gv±wlhbooZjZdl_jgucijbagZd Ijbf_qZl_evgh qlh m ihjZ`_gguo wda_fieyjh\ 445 \k_o\b^h\bf_xsboijZ\bevgu_]mklu_emqbhgbkhojZgyxlky \h \k_o ieZ\gbdZo ihwlhfm fu kqblZ_f ieZ\gbdh\u_ emqb lZdkhghfbq_kdbf ijbagZdhf b bo hlkmlkl\b_g_fh`_lkem`blvijbagZdhfZghfZevghklb Hq_\b^ghwlhk\yaZghkl_fqlhemqbkhklhylba[he__ ijhqgh]h obfbq_kdh]h \_s_kl\Z q_f ieZklbgdZ ieZ\gbdZ'XYHU6DODWLZdb_hldehg_gby\klj_qZxlky m\b^h\ h[eZ^Zxsbo ]mklufb fgh]hqbke_ggufb emqZfb\ieZ\gbdZoLeptosagitta ocis sp. n.; L. exigua sp. n.; L. icis sp. n.; L. nudata Kassatkina, 1973; L. collariata Kassatkina, 1973; L. uschakovi Kassatkina, 1973; L. eris Kassatkina et Selivanova, 2003; Aidanosagitta scarlatoi Kassatkina, 1971; A. macilenta Kassatkina, 1971; A. acus Kassatkina et Selivanova, 2003; A. pilum Kassatkina et Selivanova, 2003; A. murex Kassatkina et Selivanova, 2003; A. bella Kassatkina et Selivanova, 2003; A. venusta Kassatkina et Selivanova, 2003; Parasagitta septicoela Kassatkina, JZajmr_gb_ djZ_\hc f_f[jZgu m \b^h\ g_ bf_xsbo ]mkluo emq_c \ ieZ\gbdZo ijb\h^bl d lhfm qlh rbjhdbc ieZ\gbd \u]ey^bl dZd madZy ihehkdZ beb kh\k_fbkq_aZ_lWlh\klj_qZ_lkym\b^h\bajh^Z2OigoradiaWDZlZd`_mL. alba Kassatkina et Selivanova, 2003. JZajmr_gb_ bklhgq_gb_ [ZaZevghc f_f[jZgu <wlhfkemqZ_s_lbgdbbam[qbdbhl^_eyxlkyhl]heh- JbkFhjnheh]bq_kdb_ZghfZebbmkZ]bll:±o\hklh\hcieZ\gbd(L. exigua sp. n.);±qZklv[hdh\h]hieZ\gbdZkih\j_`^_gghc[ZaZevghcieZklbgdhc(L. icis sp. n.)<±]heh\ZbqZklvlmeh\bsZgZ]heh\_iheghklvxhlkmlkl\m_lam[gh_\hhjm`_gb_ZmqZklhddbrdb bf__lih\j_`^_gby(P. septicoela)=±]heh\ZbqZklvlmeh\bsZgZ]heh\_iheghklvxkhojZgy_lkyam[gh_\hhjm`_gb_ZmqZklhddbrdb bf__lih\j_`^_gby(L. exigua sp. n.)> ±qZklvlmeh\bsgh]hhl^_eZkZghfZevgufmqZkldhfZev\_heyjghcldZgb(L. icis sp. n.). ;BHEH=BY FHJY lhf 30 6 2004 D:K:LDBG:K?J=??< 446 \uBgh]^Zhgbkh\k_fhllhj]Zxlky\lZdhfkemqZ_gZ[ex^Z_lky kljZggZy ^ey ij_^klZ\bl_e_c lbiZ &KDHWR gnaWKDdZjlbgZeukZy]heh\Z[_aam[gh]h\hhjm`_gby jbk<Ijbf_qZl_evghqlhfurpu]heh\uijbwlhf khojZgyxlky \ hlebqb_ gZijbf_j hl wda_fieyjh\ ba ]_hnbabq_kdb Zdlb\gh]h jZchgZ [ DjZl_jghc LZdb_ ZghfZevgu_ wda_fieyju h[gZjm`_gu kj_^b jh^h\ Leptosagitta (L. exigua sp. n., L. icis sp. n.) b Parasagitta (P. septicoela). JZajmr_gb_ dbr_qgh]h wibl_eby jbk < = Ba\_klgh qlh mfhjkdbo klj_ehd \ jZchgZo k \ukhdhc ]_hnbabq_kdhc Zdlb\ghklvx iheghklvx khojZgyxlky ldZgb dbr_qgh]h b ihdjh\gh]h wibl_eby DZkZldbgZ JZajmr_gb_ dbr_qgh]h wibl_eby m fhjkdbo klj_ehdg_hlf_q_ghgb\h^ghfbawdheh]bq_kdbg_[eZ]hihemqguo j_]bhgh\ F_oZgbq_kdb_ ih\j_`^_gby dbrdb ijb jZa[hj_ ijh[u \hafh`gu ebrv \ lhf kemqZ__kebkbevghk`Zlvibgp_lhfl_eh`b\hlgh]hh^gZdh ijb wlhf ih\j_`^Zxlky b gZjm`gu_ ldZgb < [QZ`fZh[gZjm`_guwda_fieyjumdhlhjuoijbiheghc khojZgghklb gZjm`guo ldZg_c [ue ih\j_`^_g dbr_qgbd(P. septicoela, L. exigua sp. n.). JZajmr_gb_ Zev\_heyjghc ldZgb qZklv de_lhd dhlhjhc hl^_ey_lky ijb wlhf djZy Zev\_heyjghc h[hehqdb klZgh\ylky g_ jh\gufb Z dZd [u bat_^_ggufb jbk>LZdb_hldehg_gby\klj_qZxlkym\b^h\h[eZ^ZxsboohjhrhjZa\blhcZev\_heyjghcldZgvxLeptosagitta ocis sp. n.; L. exigua sp. n.; L. icis sp. n.; L. nudata Kassatkina, 1973; L. collariata Kassatkina, 1973; L. uschakovi Kassatkina, 1973; L. eris Kassatkina et Selivanova, 2003; Aidanosagitta scarlatoi Kassatkina, 1971; A. macilenta Kassatkina, 1971; A. acus Kassatkina et Selivanova, 2003; A. pilum Kassatkina et Selivanova, 2003; A. murex Kassatkina et Selivanova, 2003; A. bella Kassatkina et Selivanova, 2003; A. venusta Kassatkina et Selivanova, 2003. :\lhju [eZ]h^Zjyl <: =hjyq_\Z <D KZ\q_gdh bH<IhghfZj_\ZaZihfhsvijbhl[hj_ijh[ KIBKHDEBL?J:LMJU :klZoh\ :K Eblheh]by hkZ^dh\ fZl_jbdh\hc hdjZbgu \hklhdZ:abb<eZ^b\hklhd>ZevgZmdZk ;_eZg L: Wdheh]bq_kdb_ bkke_^h\Zgby \ aZeb\_ Klj_ehd b [mol_Jbnh\hc =b^jhf_l_hjheh]bq_kdb_ijhp_kkugZ r_evn_ hp_gdZ \ha^_ckl\by gZ fhjkdmx kj_^m <eZ^b\hklhd>ZevgZmdZK± DZkZldbgZ :I S_lbgdhq_exklgu_ fhj_c KKKJ b khij_^_evguo\h^EGZmdZK± DZkZldbgZ:I FZkkh\u_ ZghfZebb \ ieZgdlhg_ hdSZbgguo fhS_cbkhiS_^_evguo\h^Lboh]hhd_ZgZ>hdeJ:G LK± DZkZldbgZ:IIZgl_e__\ZEGLjb gh\uo fZkkh\uo \b^Z fhjkdboklj_ehdk_f_ckl\Z6DJLWWLDGDHKKDHWRJQDWKDba Yihgkdh]h fhjy Ahhe `mjg L K 1238–1245. DZkZldbgZ:I, K_eb\Zgh\Z?GGh\u_\b^ufhjkdboklj_ehd KKDHWRJQDWKD jh^h\ Oligoradiata, Leptosagitta b Mesosagitta ba [molu Db_\dZ Yihgkdh]h fhjy ;bhe fhjyZLK± DZkZldbgZ :I, K_eb\Zgh\Z ?G KhklZ\ jh^Z :LGDQRVDJLWWD KKDHWRJQDWKD khibkZgb_f gh\uo \b^h\ ba f_edh\h^guo [mol k_\_jhaZiZ^ghc qZklb Yihgkdh]h fhjy ;bhefhjy[LK± K_j]__\ :N =hjyq_\ <: Khcn_j <G b ^j Bkke_^h\Zgb_ jZkijhkljZg_gby jZ^bhgmdeb^h\ b ly`_euo f_lZeeh\ \ [mol_QZ`fZbijbe_]Zxs_cZd\ZlhjbbaZeb\ZKlj_ehd Yihgkdh_fhj_L_a^hde9>Zevg_\hkldhgnihaZih\_^ghfm ^_em ± hdly[jy ] <eZ^b\hklhd >ZevgZmdZK 252–253. Kb\bgp_\X<<ukhpdbc<E>Zgbeyg<:JZ^bhwdheh]bq_kdb_ ihke_^kl\by jZ^bZpbhgghc Z\Zjbb gZ Zlhfghc ih^\h^ghc eh^d_ \ [mol_ QZ`fZ :lhfgZy wg_j]by L\uiK± Duvert M., Salat C. Ultrastructural studies on the fins of chaetognaths // Tissue Cell. 1991. Vol. 22. P. 853–863. Ghirardelli E. 6RPHDVSHFWVRIWKHELRORJ\RIWKHKKDHWRJQDWKD Adv. Mar. Biol. 1968. Vol. 6. P. 271–375. Kassatkina A.P., Goryachev V.A., Sergeev A.F. Unusual morphological anomalies of Chaetognatha in planktonic ecosystem from Chazhma Bay of Japan Sea // CREAMS-2000. Proc. Int. Symp. on Oceanography of the East Asian Marginal Seas. Vladivostok. 2000. P. 28. Kassatkina A.P., Pantelyev E.V., Yurasov G.I. et al. MorphologiFDODQRPDOLHVRIKKDHWRJQDWKDLQ3HWHUWKH*UHDW%D\RIWKH Sea of Japan // Oceanography of the Japan Sea. Vladivostok: Dalnauka. 2001. P. 255–259. Sergeev A.F., Goryachev V.A., Soyfer V.N. et al. The oceanography aspect of radiation situation in Chazhma Bay of Japan Sea // CREAMS-2000. Proc. Int. Symp. on Oceanography of the East Asian Marginal Seas. Vladivostok. 2000. P. 39. ;BHEH=BY FHJY lhf 30 6 2004
